I mean that since you converted the strokes to path it's no longer one segment.
Rather, now the previous visual area of the stroke is built up from a path with two sides.
You unconnected four nodes at the top.
If you add a stroke and no fill to the path you can see that now on both of the previous segments ends are unconnected.
The black fill used in your open path now renders as a bubble membrane, filling up the space in between.